Name Cr Method of study Time Location Organiser
Evolutionary Biology- book re-exam Futuyma & Kirkpatrick 5 Cr Examinarium (electronic exam room) 20.12.2019 - 31.7.2020
Evolutionary Biology 10 Cr Examinarium (electronic exam room) 19.10.2019 - 13.12.2019
Evolutionary biology 10 Cr Course 9.9.2019 - 25.10.2019
Evolutionary Biology 10 Cr General Examination 21.12.2018 - 21.12.2018
Evolutionary Biology 10 Cr General Examination 23.11.2018 - 23.11.2018
Evolutionary Biology 10 Cr General Examination 26.10.2018 - 26.10.2018
Evolutionary biology 2018 10 Cr Course 24.9.2018 - 26.10.2018
Evolutionary biology 10 Cr Course 25.9.2017 - 27.10.2017

Target group

Primarily Master’s students in Ecology and Evolutionary biology.

Learning outcomes

The aim of the course is that students

  • Learn the basic concepts, theory and mechanisms of evolution from the level of genes to species.
  • Learn to perform data analysis independently and apply the evolutionary framework to interpret the results
  • Improve their skills in preparing and presenting a data analysis project
  • Improve their skills in critically reading and reporting on current literature.


1. year, 1. period.


The course consists of independent lectures (1. week, ~18 h), three intensive weeks of course work (2.-4. week, ~35 h each) and a book exam. Lectures cover general evolutionary theory supporting the course literature (Futuyma & Kirkpatrick: Evolution). The lectures and coursework will concentrate on how genes and genomes change during the course of evolution, how we can study them and what can we learn from these studies. During the intensive weeks, students do practical exercises, data analysis and seminar work, concentrating on one topic each week: i) population genetics, ii) phenotypic evolution and iii) molecular evolution and speciation.


Attendance is compulsory during weeks 2-4. The intensive weeks include introductory lectures, exercises, data analysis and seminar work. During these weeks students also read scientific papers, discuss and present their findings. Students are expected to familiarize themselves with and actively participate in discussion topics and complete and present the data analysis projects adequately.

The book exam is taken in Electronic exam rooms (Examinarium: in October-December. Separate registration in WebOodi is required for the book exam.

Study materials

Douglas J. Futuyma, Mark Kirkpatrick 2017: Evolution, 4th edition

Assessment practices and criteria

The final grade (1-5) is based on activity and performance during the lectures and course assignments (50%) and the book exam (50%). The book exam must be passed before the final grade is given and passing the book exam requires half the points available.

Relation to other study units

Replaces the former course 52333 Syventävien opintojen kirjatentit: Evolutiikka, Futuyma.